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Brain Res (2018) 1678:6474

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353 These findings confirm that in the synovium, GLP-1R is expressed on two different cell types, macrophages and fibroblast-like synoviocytes, which are specialized cells distributed within the synovial intima and subintima, and these cell types play important roles in hyaluronic acid synthesis, metabolite processing, and clearance of matrix degradation fragments

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One of these was GIP which Turner took back to Surrey University where he had moved to from the Royal Free Hospital